Research Areas
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Life Science / Biodiversity and systematics
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Environmental Science/Agriculture Science / Conservation of biological resources
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Life Science / Evolutionary biology
Graduate School
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The University of Tokyo, Graduate School, Division of Agricultural Science
Doctor's Course, 1986.03, Completed
External Career
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, Researcher, 1989.02 - 1995.03
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, Researcher, 1989.09 - 1990.03
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, Researcher, 1995.04 - 1997.03
Papers
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A new species of nine-spined stickleback, Pungitius modestus (Gasterosteiformes, Gasterosteidae), from northern Honshu, Japan., Zootaxa, 5005(1) 1-20, 2021.07
Matsumoto T, Matsuura K and Hanzawa N.
Multiple Authorship (Only Japanese)
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Anew silverside, Doboatherina palauensis (Atheriniformes: Atherinidae) from the Palau Islands., Ichthyological Research, 68(1) , 2020.08
Kimura S, Takeda K, Gotoh RO and Hanzawa N.
Multiple Authorship (Only Japanese)
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Genetic structure of glacial-relict populations of a freshwater sculpin, Cottus nozawae, in northeastern Japan., Biogeography, 20 96-102, 2018.09
Ito N, Gotoh OR, Shirakuma T, Araki Y and Hanzawa N.
Multiple Authorship (Including Foreigners)
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Occurrence of Eugymnanthia sp. (Leptmedusae, Eirenidae), an evolutionary advanced species of the bibalve-inahbiting Hydrozoans, from Ongael Lake, Palau Islands, associated with Brachidontes sp. (Bivalves, Mytilidae)., Kuroshio Biosphere, 14 22-26, 2018.01
Kubota S. and Hanzawa N.
Multiple Authorship (Including Foreigners)
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Phylogenetic relationships and morphological variations of upside-down jellyfishes, Cassiopea spp. inhabiting Palau Islands., Biogeography, 19 133-141, 2017.09
Arai Y, Gotoh RO, Yokoyama J, Sato C, Okuizumi K and Hanzawa N.
Multiple Authorship (Including Foreigners)
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Distribution and dispersal of the giant jellyfish Nemopilema nomurai (Scyphozoa, Cnidaria) inferred from its genetic population structure., Biogeography, 19 35-42, 2017.09
Gotoh RO, Chang S, Qu X, Ito C, Okuizumi K, Yoon W, Iizumi H and Hanzawa N.
Multiple Authorship (Including Foreigners)
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Two new species of Jesogammarus from Japan (Crustacea, Amphipoda, Anisogammaridae), with comments on the validity of the subgenera Jesogammarus and Annanogammarus., Zoosystematics and Evolution, 93(2) 189-210, 2017.03
Tomikawa K, Nakano T and Hanzawa N.
Multiple Authorship (Including Foreigners)
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Molecular systematics and DNA barcoding of Altai osmans, oreoleuciscus (pisces, cyprinidae, and leuciscinae), and their nearest relatives, inferred from sequences of cytochrome b (Cytb), cytochrome oxidase c (Co-1), and complete mitochondrial genome., Mitochondrial DNA Part A, 2016.03
Kartavtsev Y, Batischeva NM, Bogutskaya NG, Katugina AO and Hanzawa N.
Multiple Authorship (Including Foreigners)
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Collaboration between Yamagata University and Kamo Aquarium - Rapid adaptive evolution of jellyfishes inhabiting marine lakes in Palau., Der Zoologische Garten, 85 34-41, 2016.01
Hanzawa, N., R. O. Gotoh, Y. Nakauchi, A. Funahashi, H. Kanematsu, M. Miyashita, Y. Arai, K. Okuizumi
Multiple Authorship (Including Foreigners)
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Phylogenetic relationships of Russian far eastern flatfish (Pleuronectiformes, Pleuronectidae) based on two mitochondrial gene sequences, Co-1 and Cyt-b, with inferences in order phylogeny using complete mitogenome data., Mitochondrial DNA, 27(1) 667-678, 2016
Kartavtsev Y, Sharina SN, Saitoh K, Imoto J, Hanzawa N, and Redin AD.
Multiple Authorship (Including Foreigners)
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Rhopilema asamushi Uchida (Cnidaria, Scyphozoa, Rhizostomidae) newly occurred in Tanabe Bay, Wakayama Prefecture, Japan., Biogeography, 17 125-127, 2015.09
Hanzawa, N., R. O. Gotoh, M. Kawamura and S. Kubota
Multiple Authorship (Including Foreigners)
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Polygenic expression of teratozoospermia and normal fertility in B10.MOL-TEN1 mouse Strain, Congenital Anomalies, 55(92) 98, 2015.04
Hirawatari,K., N. Hanzawa, Kuwahara, Aoyama, I. Miura, S. Wakana and H. Gotoh
Multiple Authorship (Including Foreigners)
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A cascade of epistatic interactions regulating teratozoospermia in mice., Mammalian Genome, 26 248-256, 2015.03
Hirawatari,K., N. Hanzawa, I. Miura, S. Wakana and H. Gotoh
Multiple Authorship (Including Foreigners)
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Geographical distribution and genetic diversity of Gymnogobius sp. "Chokai-endemic species" (Perciformes: Gobiidae), Icthyological Research, 62 156-162, 2015.01
Chiba N. S., R. Kakehashi, K. Shibukawa, T. Mukai, Y. Suzuki and N. Hanzawa
Multiple Authorship (Including Foreigners)
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Genetic population structure and dispersal of giant jellyfish, Nemopilema nomurai, 61(2) 6, 2014.08
Naoto Hanzawa
Single Author
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Phylogenetic relationships of Russian Far Eastern flatfish (Pleuronectiformes, Pleuronectidae) based on two mitochondrial gene sequences, Co-1 and Cyt-b, with inferences in Order phylogeny using complete mitogenome data, Mitochondrial DNA, 2014.05
Kartavtsev, Y. P., S. N. Sharina, K. Saitoh, J. M. Imoto, N. Hanzawa and A. D. Redin
Multiple Authorship (Including Foreigners)
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Characterization of comparative genome-derived simple sequence repeats for acanthopterygian fishes., Molecular Ecology Resources, 13 461-472, 2013.04
Gotoh, R. O., S. Tamate, J. Yokoyama, H. B. Tamate and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Phylogeny and biogeography of highly diverged freshwater fish species (Leuciscinae, Cyprinidae, Teleostei) inferred from mitochondrial genome analysis., Gene, 514 112-124, 2013.01
Imoto, J., K. Saitoh, T. Sasaki, T. Yonezawa, J. Aadachi, Y. P. Kartavtsev, M. Miya, M. Nishida and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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QTL mapping of sperm-head abnormality genes causing male sub-fertility on mouse Chromosome 1 and 4., Mammalian Genome, 2012.03
Goto, H., K. Hirawatari, N. Hanzawa, I. Miura and S. Wakana
Multiple Authorship (Including Foreigners)
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Population genetic structure of the striped silverside, Atherinomorus endrachtensis (Atherinidae, Atheriniformes, Teleostei), inhabiting marine lakes and adjacent lagoons in Palau: marine lakes are "Islands" for marine species., Genes and Genetic Systems, 86 325-337, 2011.10
Gotoh, R. O., Chiba, S. N., Goto, T. V., Tamate H. B. and Hanzawa, N.
Multiple Authorship (Including Foreigners)
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Characterization of marine X-family DNA polymerase and comparative analysis of base excision repair proteins., Biochem. Biophys. Res. Com., 415 193-199, 2011.10
Kodera, H., R. Takeuchi, Y. Uchiyama, Y. Takakusagi, K. Iwabata, H. Miwa, N. Hanzawa, F. Sugawara and K. Sakaguchi
Multiple Authorship (Including Foreigners)
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Phylogenetic characterization of three morphs of mussels (Bivalvia, Mytilidae) inhabiting isolated marine environments in Palau Islands. , Zoological Science, 28(8) 568-579, 2011.08
Goto, T. V., H. B. Tamate and N. Hanzawa
Multiple Authorship (Including Foreigners)
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Species diversity and community structure of pelagic copepods in the marine lakes of Palau. , Hydrobiologia, 666(1) 85-97, 2011.05
Saitoh, S., H. Suzuki, N. Hanzawa and H. B. Tamate
Multiple Authorship (Including Foreigners)
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Preliminary review and conservation of Gymnogobius castaneus complex in Japan., 57(2) 173-176, 2010.11
Mukai, T., K. Shibukawa, T. Shinozaki, H. Sugiyama, S. N. Chiba and N. Hanzawa
Multiple Authorship (Only Japanese)
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Phylogenetic characterization of marine bacteria that induce morphogenesis of the red alga, Porphyra yezoensis. , Fish Genetics and Breeding Science , 40(1) 29-35, 2010.09
Hanzawa, N., S. N. Chiba, S. Miyajima, A. Yamazaki and N. Saga
Multiple Authorship (Including Foreigners)
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Comparative genome analysis of mammalian NKG2D ligand family genes provides insights into their origin and evolution. , Immunogenetics, 62 441-450, 2010.07
Kondo, M., T. Maruoka, N. Otsuka, J. Kasamatsu, K. Fugo, N. Hanzawa and M.Kasahara
Multiple Authorship (Including Foreigners)
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Molecular phylogenetics of pricklebacks and other perciod fishes from the Sea of Japan. , Aquat. Biol., 8 95-103, 2009.12
Kartavtsev, Y. P. S. N. Sharina, T. Goto, O. A. Rutenko, V. V. Zemnkhov, A. A. Semenchenko and N. Hanzawa
Multiple Authorship (Including Foreigners)
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Peripatric differentiation among adjacent marine lake and lagoon populations of a coastal fish, Sphaeramia orbicularis (Apogonidae,Percifirmes, Teleostei)., Genes and Genetic Systems, 84(4) 287-295, 2009.10
Gotoh, R. O., H. Sekimoto, S. N. Chiba and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Comprehensive phylogeny of the family Sparidae (Perciformes: Teleostei) inferred from mitochondrial gene analyses., Genes and Genetic Systems, 84 153-170, 2009.06
Chiba, N. S., Y. Iwatsuki, T. Yoshino and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Sequence diversity at cytochrome oxidase I (COI) gene among sculpins (Scorpaeniforms, Cottidae) and some other scorpionfish of Russia Far East with phylogenetic and taxonomic insights. , Genes and Genomics, 31(2) 183-197, 2009.06
Kartavtsev, Y. P. S. N. Sharina, T. Goto, A. A. Balanov and N. Hanzawa
Multiple Authorship (Including Foreigners)
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Sequence diversity at cytochrome oxidase I (CO I) gene among sculpins (Scorpaeniforms, Cottidae) and some other scorpionfish of Russia Far East with phylogenetic and taxonomic insights.(共著), Genes and Genomics, 21(2) 183-197, 2009.04
Kartavtsev, Y. P., S. N. Sharina, T. Goto, A. A. Balanov and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Cytochrome oxidase I gene sequences analysis in six flatfish species (Teleostei, Pleuronectidae) of Far East Russia with inferences in phylogeny and taxonomy.(共著), Mitochondrial DNA, 19 479-489, 2008.12
Kartavtsev, Y. P., S. N. Sharina, T. Goto, A. N. Chichvarkhin, A. A. Balanov, K. A. Vinnikov, V. N. Ivankov and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Inferences in Leucuscinae (Pisces, Cyprinidae) phylogeny and taxonomy based on cytochrome b sequence distances and on enzyme loci diversity.(共著), Kor. J. Genetics, 29(4) 427-435, 2007.12
Kartavtsev, Y. P. and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Phylogeographic origin of Hokkaido house mice (Mus musculus) as indicated by genetic markers with maternal, paternal and biparental inheritance., Heredity, 96 128-138, 2006.01
Terashima, M., S. Furusawa, N. Hanzawa, K. Tsuchiya, A. Suyanto, K. Moriwaki, H. Yonekawa and H. Suzuki.
Multiple Authorship (Including Foreigners)
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Allozyme and morphometric research of two common mussel species of genus Mytilus (Mollusca, Mytilidae) in Korea, Japan and Russian waters., Kor. J. Genet., 27(4) 289-306, 2005.12
Kartavtsev, Y. P., A. Y. Chichvarkhin, A. Kijima, N. Hanzawa and I.-S. Park.
Multiple Authorship (Including Foreigners)
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Genetic variability of MHC class II β1 genes in Far Eastern dace, Tribolodon hakonensis. , Mar. Biotech., (6) 398-403, 2004.09
Kasamatsu, J., Y. Watanabe and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Genetic divergence of Far-Eastern dace species belonging to the genus Tribolodon (Pisces, Cyprinidae) and closely related taxa., Russ. J. Genet., 38(11) 1285-1297, 2002.01
Kartavtsev, Y. P., V. V. Sviridov, N. Hanzawa and T. Sasaki.
Multiple Authorship (Including Foreigners)
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Genetic differetiation and phylogenetic relationships of cyprinid fishes, genera Tribolodon and Pseudaspius inferred from mtDNA sequences., Proc. Int'l Conference of Evolution, Genetics, Ecology, and Biodiversity., 8, 2001.10
Hanzawa, N., T. Sasaki, T. Uematsu, Y. P. Kartavtsev, and V. V. Sviridov.
Multiple Authorship (Including Foreigners)
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Evolutionary features of a cyprinid fish, genus Tribolodon in the Far East inferred from molecular markers. , Proc. Int'l Meetings of Biodiversity in Asia 2000., 25, 2000.09
Hanzawa, N., T. Sasaki, V. V. Sviridov, and Y. P. Kartavtsev.
Multiple Authorship (Including Foreigners)
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Factors in growth and morphogenesis recovery under axenic conditions in Porphyra yezoensis (Bangiales, Rhodophyta), especially the effect on symbiotic bacteria in conditioned media., Bull. Inst. Oceanic Res. & Develop., Tokai Univ., (21) 57-76, 2000.04
半澤 直人
Multiple Authorship (Including Foreigners)
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16SrDNA-based phylogenetic analysis of marine flavobacteria that induce algal morphogenesis., J. Marine Biotech., 6(2) 80-82, 1998.01
Hanzawa, N., K. Nakanishi, M. Nishijima, N. Saga.
Multiple Authorship (Including Foreigners)
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16S rDNA-based phylogenetic analysis of marine flavobacteria., J. Marine Biotech. , 3(1-3) 111-114, 1995.10
Hanzawa, N., S. Kanai, A. Katsuta, Y. Nakagawa, and K. Yamasato.
Multiple Authorship (Including Foreigners)
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Heavy oil degrading bacteria isolated by long-term enrichment with almina colums containing heavy oil C., Biosci. Biotech. Biochem., 59(11) 2159-2161, 1995.09
Shirai,K., N. Hanzawa, and A. Katsuta.
Multiple Authorship (Including Foreigners)
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Growth requirements of hyperthermophilic sulfur-dependent heterotrophic archaea isolated from a shallow submarine geothermal system with reference to their essential amino acids., Appl. Environ. Microbiol., 60(8) 2898-2904, 1994.08
Hoaki, T., M. Nishijima, M. Kato, K. Adachi, S. Mizobuchi, N. Hanzawa, and T. Maruyama.
Multiple Authorship (Including Foreigners)
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Epidermal growth factor stimulates protein synthesis in primary cultures of salmon hepatocytes., Gen. Comp. Endocri., 90 383-386, 1993.09
Duan, C., Y. Takeuchi, N. Hanzawa, and S. Miyachi.
Multiple Authorship (Including Foreigners)
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Isolation and characterization of a marine bacterium producing a seed germination inhibitor., J. Marine Biotech., 1(2) 105-108, 1993.09
Hanzawa, N., F. L. Singleton, E. Hamada, R. R. Colwell, and S. Miyachi.
Multiple Authorship (Including Foreigners)
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Use of primary cultures of salmon hepatocytes for the study of hormonal regulation of insulin-like growth factor I expression in vitro., Zool. Sci., 10(3) 473-480, 1993.06
Duan, C., N. Hanzawa, Y. Takeuchi, E. Hamada, S. Miyachi, and T. Hirano.
Multiple Authorship (Including Foreigners)
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Mitochondrial DNA polymorphisms in subterranean mole-rats of the Spalax ehrenbergi superspecies in Israel, and its peripheral isolates., Mol. Biol. Evol., 10(3) 590-604, 1993.04
Nevo, E., R. L. Honeycutt, H. Yonekawa, K. Nelson, and N. Hanzawa.
Multiple Authorship (Including Foreigners)
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Collection of marine bacteria., MBI Report, 1992 36-37, 1992.03
Hamada, E., N. Hanzawa, S. Kanai, and A. Katsuta.
Multiple Authorship (Including Foreigners)
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Genetic control of sex-dependent meiotic recombination in the major histocompatibility complex of the mouse., EMBO. J., 10(3) 681-686, 1991.04
Shiroishi, T., T. Sagai, N. Hanzawa, H. Gotoh, and K. Moriwaki.
Multiple Authorship (Including Foreigners)
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Recombinational hotspot specific to female meiosis in the mouse major histocompatibility complex., Immunogenetics, 31 79-88, 1990.04
Shiroishi, T., N. Hanzawa, T. Sagai, M. Ishiura, T. Gojobori, M. Steinmetz, and K. Moriwaki.
Multiple Authorship (Including Foreigners)
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Geographical differentiation in populations of Japanese dace Tribolodon hakonensis deduced from allozymic variation., Zool. Sci., 5(2) 449-461, 1988.04
Hanzawa, N., N. Taniguchi, and K. Numachi.
Multiple Authorship (Including Foreigners)
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Variability of mitochondrial DNA in Japanese dace, Tribolodon hakonensis (Cyprinidae)., Jpn. J. Genet., 62(1) 27-38, 1987.02
Hanzawa, N., H. Yonekawa, and K. Numachi.
Multiple Authorship (Including Foreigners)
Books
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RED DATA BOOK YAMAGATA 2019., 2019.03
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Sources and dispersal of the giant jellyfish Nemopilema nomurai inferred from its genetic population structure, 2017.10
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Genetic diversity and evolution of marine animals isolated in marine lakes. Chapter 7 in Analysis of Genetic Variation in Animals., InTech, 2012.02
Hanzawa, N., R. O. Gotoh, H., Sekimoto, T. V. Goto, S. N. Chiba, K. Kuriiwa and H. B. Tamate
Other research activities
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Red Data Book Yamagata-Freshwater Fish
Fieldwork
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Isolation, Cultivation, Characterization, and Preservation of Marine Bacteria
Fieldwork
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A Survey of Freshwater Fishes of Kochi Prefecture
Fieldwork
Grant-in-Aid for Scientific Research
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Grant-in-Aid for Scientific Research(C),2023.04 - 2024.03
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Grant-in-Aid for Scientific Research(C),2011.04 - 2014.03
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Grant-in-Aid for Scientific Research(C),2005.04 - 2008.03
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Grant-in-Aid for Scientific Research(B),2004.04 - 2007.03
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Grant-in-Aid for Scientific Research(C),1999.04 - 2003.03
Presentations
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The 5th International Jellyfish Workshop, International presentation, 2008.12, Oral presentation (general)